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Aglais io, Linnaeus (1758)

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Infraspecifics
acilia myrrha versicolor isobeon bellona selene lineata gongylodes sela camillus nivescens westringi trebula catops crocophala eumedon porus cassander ripartii eff eupheme argus pieroides hainani cosyra cocles thersites heptagon fausti janeira klugius eurypilus cometes belti lutea turcicus spinipes miris memusae albisparsa pylaon ducalis maenalis violetae bituberculata quinquepartitus contempta diores anona sephirus juncta curvilinea nivea phlaeas belfragei regalis leucane zephyrinus glaphyra illustris thyodamas elaborata ocellatus splendens noureddin dentifasciata barbotini nicaea io tabulata hoffmannsi viridescens homogena megacephalum komarovi larra cicatricosa diadema apollinairei odana brookiana verbasci caja metzli strigata alba aegeria trojana festiva naranja phthisica aliris melanura festiva zozine berania anadyomene jeskovi lathyi umbra nupta elpenor egeus chiron hyperbius directa xiphioides busiris exprimens hippophaes bharata crethon pandora argiopiformis fenestraria cilisca cyanocephalus phoebus athamas eleuchea paphia cornutus caecutiens aurelius treitschkei marudhamneydhal apollo axifera furcula sagana patagiatus apta amethystus fimbria ponticeriana margelanica fusca harmonia taeniolatus sclopetarius hypermnestra darius viridis daudini virgaureae splendens orsilochus dewysea fusiformis discrepans martia leonina marshalli populi utricularis petreus nigrina arcuata materna atreus rosea moyeri taraxaci minutum zerynthia belladonna acalypha homaena engeli spectrum sardanapalus leucocephalum marcella berinda gibberosa tyrannus eurilochus liltvedi spinosus huttoni muscorum lativitta maculata caerulea telamonius papillosa temporalis intercalaris septica placida latipes memnon colemani tilburyi ferghana fragilis adamsi sanaca spiculata olivacea oedipus arianeae chapmanorum wagneri psittacinum arvensis pasithoe bisaccata violacea uranus japonica aymonieri wagneri cinereum auratus hutchinsoni virginica illioneus spectans pennocki convolvuli polycephalum auriculatus aganippe phrynosoma micans obscura pustulosus afghana graminea auronitens belisama lemniscata pilumnus aestuans brasiliensis donata Largen & Drewes gurkoi cuetzpali avinovi harpalyce gabonensis semiaurata sonorina teucer caranea macrops lichenigera lythrochila zebuda zilloides hypothous caffra lituraria boulengeri proserpina mixteca agostina arizonica nerii iris seleucida eriophora conifera beanii montecristoi arcensis periboea foxi placida aequabilis reevesii subsignata africana mongolica moreletii incrassata torenia virgo automedon inangulata anguineus maculosa asio bessarabicus argenthona labyrinthea splendens cerialis feae strigulosa blainvillii blaptoides eucharis funebris adramitanus japonica cassiope kharini centralasiae braconnieri cancellatus hyparete furcata atricollis xanthostoma xanthus magdala gabonica euphorbiae brevirostris clatratus gracilis branchi atrata sacrifica capillata cornutum catenulatus lucasi gregorii haemorrhoidalis batea mariamne millsae spectabilis coronatum concolor mysis mitrata ugandaensis geryon howardi norvegica incurvata douglasii convexus australis sagittata aculeata cornelia philaretes agricola implexa histrio hernandesi coriaceus hirsutus sanctispiritus africana amata eryx giganteus mcallii creutzeri sambawana schreibersi agama angustipennis bispiralis plagiata modestum excellens glandon swainsoni anchietae dimidiata tubularia karelini orbiculare faustus podarce triserrata armata intermedia avernus promethea columbiana quadripunctaria finitimus orbitulus clymene atra maculata securifera vermicularis reversa platyrhinos galicianus brevirostris inaurata bibronii samarcandica philoctetes angulifera spirorbis reversa solare gebleri heinsohni clavipes gracilimembris syriaca ulei galathea enigmaticus clymene taurus gigas antiopa plumipes hartmanni bernardoespinozai gerdrudtus helicina dominula marsyas glabratus californica pilipes kirkii herois helenor lachesis rangii quadripunctaria damo granulatus cyanomelas clavipes lebretoni crinis peleides larissa annulatus splendidior arota gyllenhali polychloros clavata lionotus macrops antimachus titea venustissimus lecontei dione henningi vaualbum edulis mossambica ephesperis epistrophus russiae chinensis argus editha hispanus xanthomelas fenestrata mwanzae albicinctus occitanica subpunctatus slovenica gorgon hortensis albissima inaurata persimilis achilles ines sagittarius striata xanthoides hummeli ancocisconensis plumipes picticauda subochrea arge mucosa bipunctata heteronea insulicola arenicola borbonica planiceps phaeton epimede korros albipuncta rubidus intricatus bellissima cruentata rueppelli maturna deidamia semicarinatus cruentata hyllus irregularis campestris livida sankaranica cynthia hanria dhumnades elata albicans jankowskii decemnotata queribunda tassiliensis iduna ariadne pteria luzonensis ornatrix bellargus lafossei denverensis venustula weidholzi gillettii merione bicolor longifrenis pectinata coridon lineatus depressula tsushimensis laungwalaensis anicia dryope zambesiaca nebulatus pulchella corydonius linnaei duodecimguttata bistriata subcristatus chalcedona ophione angulata atra pulchelloides hispana lopatini formosa dehaani brevicauda editha alida fasciata salomonis arcania loschnikovi fulgida hupingensis cypriaca aurinia dorcas scabra sumatrana gardetta lusitanicus hirticollis graeffei sacra desfontainii bungaroides maeander hybrida bradleyi stellio lauta teleboas pygela stephensii stellatarum marginalis lengi eburnus tuberculata acerosa orites nigrescens dorus maurus limbalis australasia vulgaris volupe salambria thwaitesi bicolor melanostigma glycerion monilis longilabris mouhoti himalayana besanti aconitus gana semenovi postrubida leander montivagus nigrior nitidula lehmanni fournieri numilia litigiosa albolabris alecto oedippus morbillosus ohlone maudae stoliczkana mimosae japetus insularis pamphilus nemoralis oregona brevispina caucasia mittrei aglaura flesus atlantica symphita nitens patruela nordviei arabicus fumosa hewitsonii nestus berryi thyrsis olympiae pugetana arenata helioscopus semirosea parra carolinensis tullia pedemontanus punctulata furcifera horvathi dissimilis boisduvali tethys aquaticus archippus piochardi purpurea tauricornis interscapularis canicosta davidii sikorae arthemis problematicus repanda sia maculatus leucitys sophonisba fimbriatus scutellaris prina mystaceus Epiphora tatila kobela desiccata vulgaris regalis sexguttata crassa sakoi bauhiniae chlorochroa indrani lobata lorquini rugosus splendida trinotata scutellatus atlas pusilla pinsbukana gorochovi ichnusa weidemeyerii rumelicus sylvatica diodia versicolor caesar monima agni coronatus camilla rutilans sylvicola atrica vlangalii erebus concordia denuba coronatoides caschmirensis andromica scabrosus tenuicincta x-notata dhofarensis taprobanis clytia diabolica ladakensis diaphanus scheidleri theatina acco jensvindumi alcinoe norica brucei milberti libethris smaragdinus tranquebarica acdestis sinaitus aslauga margarita lucetia ellinorae rizana morgane solieri coerulea actius agilis angulosa eburnea celebica minimum urticae oto splendens apollonius agnetae flavistriga macris phisara maasi gannascus theudelinda stscheglowi duponti ariadne flavimaculatus nigridorsis bhagava morsomus cyananthe pireta duponti autocrator mutabilis oblonga amphinome sinica sanguineus cyaniris andromeda aurofasciata behrii ruderatus ocellata arete setosum ethusa confusa nevadica boedromius sanguinolentus thrasibulus arethusa savignyi orsis estella torosus bremeri savignii enega arinome rara alberti hyperia flora truncaticollis clypeata cardinal batillifera belladonna formosana richardi ilithyia gunilla fallax charltonius taylori toxopei chloe sambara pelagicus procilla onega ullrichii manga cephalus coendersi ardoris epinome pygmaea athemon rubescens vagans binotata clodius insolitus volcanica februa affinis diaphana anna tigilla variolosus squamigera davydovi auriguttata alala longicollis astala hypaesia vinctus luizae delphius reticulata iridicolor bridgesii candrena aita violaceus dorsata epaphus elongatalis boreas feronia vasava canaliculata clymena bifasciata vladimirskyi mexicana eversmanni argentalis basiloides avicularia brachyurus eluina burnii boulengeri vittata felderi melilialis iphiclus glauconome juruensis bucephala euclides echerius antakarana expansa glacialis taphiusalis purpurea torpida neglecta fylla brunoi arquata hardwickii subfasciata bredowii guatemalena rufa grotei pandama gerontes brygooi bubalus honrathi sinuosus californica azuraklaasi combusta adippe geza confidens alta huberi pudicula eulalia iphthime caei sundana elisa neavei decaryi bisonia imperator pallida serpa laodamia minatrix flavescens kamala neophron desperata taurina jacquemontii acaciusalis cytherea velutina merianae limbata nerippe tantalus ebenaui diceros maharaja cyclothyris fessonia adrasta hirschii reticulatus niobe savitri griveaudi carinata maximinus strigigrapha syma orea taunayi darwini polyxena perroti micra tartarea mnemosyne ovifera messana laothoe geroldi deccanensis rumina egeon minima querci nomion capillatus plesaure postverta huriana superba podalirius durga nana definita simo scitaria thessalia agacles metallica brachycera feisthamelii ouida perarmata salamandra simonius fenestrella uniformis tithia urticans cornuta podalirinus eugenes peyrierasi belfragei smintheus usitata rubriplaga lynnae lepidocera acraea ramanantsoai cornutula staudingeri maculata plumbeofusa coenus variegata senegalensis agathon flegyas stumpffi excelsa stenosemus gloriosae faganaria arene bicegoi coerulea crataegi superciliaris scalaris stoliczkanus marginata ozophanes anubis claudina aurata hippia noctula tedi bimaculata stubbendorfi horsfieldii sanguinea sanguinalis lepadiformis leucodice infernalis therezieni uhleri szechenyii inachus mitigata dyonis busirus molle porcellus cecilia thieli globulare tenedius limborgii abhadraca gisella epicra moluccensis askoldensis patrona tristis caryae paralekta convergens texa ossipandae rivularis elvina tuberculata gibbera tianschanicus philarchus comptaria hydaspes eleusina picta affinis franciscana vadoni sanguineoplaga paphia obstataria astarte hewitsonius robusta aksana lycisca kushaica galeata brevicauda buxtoni punctaria pygas mithridates sabbadini alaiana nicaeus concinna merdigera indra drucei puppillaria sorana papinianus taureanensis beresowskii gnidus cuvieri cheni machaon sylvia linearia pitheas tamenund flava croatica carausius hilaris lilii polyxenes austenia ruficiliaria cyclops thraso diminuta dentata acheus olivieri asparagi zelicaon lennigiaria eunomia maia diffinis lucianus anthyllidis duodecimpunctata zalmoxis taiwana hemes picta fuciformis azanoides lavandulae macilenta nireus pendularia jovianus assimilis gracilis rhadamanthus quatordecimpunctata constantinus medon heydena jurinei lancea fulgidus ottonis balbinus oxytropis quinquepunctata dardanus hebes nanaria humboldtii cerialis aperiens radians caricae rubicundus paracenthesis phorcas xypete albipunctata arizana rubra propitia erythrus cyanella demodocus eberti hyponoea bomplandii bhaga saundersii zygia minos daturaphila demoleus themis annularia contubernalis cana staudingeri purpuralis solani oribazus inanum laeta cardases cervina thetis meneria graslini trivittata hesperus phaethusa flaveolata alicia fridericus nitobeana thysbe aulestes centaureae dorsalis lormieri cruentaria oileus recta tityus formosissima laeta ovalis menestheus purpuraria oppelii oileus sinensis venata arizonensis corsica sexpunctata mangoura preussi acmaea adepta bottae gallii borealis sarpedon erichsonii antonio alacris calabra communis umbratica muticum punctum melanopus noblei edwardsii vibicaria tryphena orcus jayakari kuldjaensis nemesis carniolica sayi demolion eleus steganioides albezens atropos ligustri virginiensis occitanica bivittata euchenor eupalus semiustus pyrame philetas styx lineolatus fulmen exulans atrata liomedon harpalyce cosmospila flavilla malvae lachesis rhacotis viciae azurea anactus neophron turbata gautama cinarae mantinea loti circumdata aegeus rex achlyobathra buddha jupei ino romeo denticollis ambrax burejana superans armoricanus kindermannii octauius osterodensis flaveola polytes dohertyi discophora brahma serratulae antarctica sylphina dorycnii margaritacea castor levana borneamagna ussheri sidae limacina licursis ephialtes nebulosa dravidarum plenorbis melotis agesilas transalpina piperata fuscus aliena vulgaris demophon alveus sula filipendulae rubiginosa helenus artemis digammata amydon onopordi pixe lonicerae viridis iswaroides chapae ocellata aidea carlinae gekko lycisca trifolii vittata nephelus dubernardi vestalis troglodyta erosus klemmeri arcius geryon vibex lowi gnoma limboundata canescens ferghana periander statices purpurata mayo groenendaeli emutaria nessus artobolevskiji croceus americana sexpunctata memnon keralana isomerica andria fides alfacariensis metallica succinea polymnestor luna immorata fabius aberrans nupta chlorocoma dimidiata punctata rumanzovia ningpoana immutata nobilis arsalte christina globulariae texana alcmenor rhodopneuma opicata glycerium ericetorum neonympha electo budensis lunifera macilentus selene lydia ryphea alana elis notata guttata taiwanus villica minorata nessus domicella gracilis erate chloros boliviana maraho aureus marginepunctata clytemnestra felderi tenuicornis pallidula agestor imperialis imitaria isidora loxus pudica fieldii volgensis subferruginea slateri semihyalina ellops polla repudiata harfordii erythropyga hectica clytia figulatum otolais pulverulenta golbecki hyale moorei clavata bianor pardaria zampa remissa meadii coa nervosus hoppo temerata risa geometrina gratiosa myrmidone kuroshimae bicolor krishna percnosticta intermedia tryxus nilagiriensis nonatoi sanguinicollis paris margarita sandace occidentalis ocellifera pluto buddha tyotaroi vestaliata timur palaeno claudia excavata crino iljai bimaculata rahrioides praeneste phicomone hegesia vittata palinurus polita c-album demophon almana xerene peranthus deva paradoxata egea manilensis atlites dorsalis montrouzieri punctata comma meleagris metelkana chorimene scapularis ulysses penicilligera prema interrogationis hispidus purpurata coenia inaequalis anchisiades bowkeri cupido c-aureum nigropunctatus sannio ethlius divaricata rosea pelaus cippus gracilis mappa amboinensis evarete imperialis andraemon lectabilis faunus stellatus pavonia celebensis genoveva viridis androgeus principalis satyrus caeruleopunctatus shadulla microlophus goudotii pentacanthum cresphontes melastigma agestis progne militaris aeolus pustulatus gregorii acrosomoides thoas crepuscularis percota bathildis weberi grisea armida okinawensis jehana caprimulgus palmyra bitias lilis hedonia ceropegia palamedes artaxerxes hieroglyphica bivexillata hebraeus ludovica hierta packardi troilus caeruela cramera walkeri sagana phaleros menophilus intermedia coccineus canadensis fellearis anteros gemmans fenestrata jason piera iphita maritatum eurymedon augusta morronensis subrepleta cayennsis sinensis lemonias keralayensis glaucus macarea hyacinthus isabellae alexis pelia crenulata neildi feredayi multicaudata iulia nicias amystis lygdamus subangulata similis nigrosuffusa higginsi rutulus ilia aurorula melanops apicalis lata oenone melanura garamas lysippus neglecta paphos boisduvalii spumans orithya scorpionoides xuthus macropoda lysippoides archidona piasus tristygma griseus pacoma cucurbitina vulnerans meliboeus bilineata chrysomelas plateada lurida rhadama displicata quadriguttata dichroa parhassus consularis concinna sophia opisthographa oxleyi theodora princeps oslari pustulifera stemosa proxima alcetas pinguis cadytis chandra peigleri ephippigera stygia branicki amyntula casta faunus daimio anacardii senatoria aegrotus terea aemula Milionia apollonia stigma karschi villida anasuja comyntas zonea aegates virginiensis viridipes zonalis appensa lacturnus fulgida smithiae iris duprei azteca morbillosus argentata minimus formosana hillapana laverna temora hamifera elegans aurantia osiris basalis metis lobesis variegatus australis tusovi buprestaria soana mexicana bufonius bruennichi cyane gavissima hortona celtis phoronea pictus catenulata sanguilineata hyalinus leilia abnormis regalis miliaris dietrichae acrocnema semialba clyton xenia sepulcralis dolichus florida aquilonaris lidderdalii idyja splendens linigera katherina bellona thaidina lancearius volcan fasciata keyserlingi chariclea leucosia alcetas walckenaeri guttula lobata dia reducta rhombicollis cornutus bicolor hedenborgii aristolochiae magnifica epithore tagis stalii curtulus rubicunda neglecta hector mascordi eunomia ausonia rhomboidea plumipes acuta pennata jophon minuta euphrosyne ausonides strumaria pigea varicolor adoxa pennicornis kotzebuea ocyaloides freija belemia variabilis orbona atratus barnesi selina mariae picta frigga crameri frontilongus butleri guianensis acamas pandiyana protensa improba charlonia luteomaculatus doxo couchii imperialis cilissa phlegon pulchella graeca penia interclusus edusa cyrtopsis masoni elima radon kriemhild aegyptiaca flavescens elegans ormondei ictis submaronica napaea falloui bilobum daplidice eques oslari lilacinus coon trifasciata pales lotta coleoptratus glauconome errans penelope lohita palu versicolor selene olympia simplex occidentalis gigas bilineata rukma rhodifer aetherea titania cecropia chloridice hammondii purpurascens schistacea neptunus minax abyssinica promethea protodice marcianus hodeva syama darwini altissima columbia sisymbrii melanogaster fenestrata vulcanus dasarada sexcuspidata thersamon annabella euryalus l-nigrum ordinoides dargei epargyros impediens excavata macrotis atalanta phidippus egerina proximus photophila natalensis latreillei brunnea usagi braziliensis virgata obtusa pulchrilatus santarosensis abnormis polyeuctes melanoviridis paweli cardui ferruginicosta radix erycina glaucippe verutum dendritica carye rutila saurita jacobaeae leucippe atrihastula proscarabaeus dimorphica sondaica submarginata scalaris jorulla inaequalis americanus gonerilla discirufa maurus horishana ixoides angusticollis hippomene lepida comma orizaba semperi nagasakiensis campanicollis indica ogina alba aurota varuna cicatrosa impressus itea cycnus nephrographa cincta cethura cressida citricola laevis kershawi euthymius monacha hesperus cardamines fasciata exanthematica violaceus myrinna aerope brunneiplaga lebeau sara lunulata hirta formosa tameamea canens dispar rectifascia scolymus maculosa moluccensis metallica terpsichore eumeus sirtalis fournierae anomalus nierstraszi unicolor regalis virginiensis marginata jasius griseus actis pustulosa rufilata vulcania sinica agrarius mungaich admetus ravilla striata doris gracilis grandis aristogiton nigromaculatus alcestis nephiloides ornata galba semicincta bernardus pavonis amandus sakiedaorum rajaei mafa menado todara borneensis scutulatus aroaniensis anastera smithi orbifer stomphax eurialus speciosus damon cepina subfusca sertorius faunula concolor harmodius spicatus daphnis cancriformis vittata alceae cambodia mathura hilaris kahruba vespertilio dorylas clavatrix consobrina tripolinus howqua oleracea marmax volans eros diardi eyndhovii helvina xylina ornata mars cerisyi escheri falcicornis imperialis camadeva dominulus nitebis deyrollei fulgens fornicata staudingeri grisea ventralis ocellatus cretica golgus geminata verhuelli lena artemis histrionica psaphon cassandra humedasae kuhli bacaenis lucia onolaus plumbalis violascens solon cerisy icarus quadrispinosa zonata luna atemeles hieroglyphica achates iphigenia sturi cecrops nereis honrathi amydra relicta argiades acilia aegeria aegeus parhassus affinis memnon agestis agestor glaphyra chlorocoma brookiana memnon alicia alida apollo xenia automedon anacardii andraemon chandra contubernalis sirtalis celtis anubis gavissima gracilis canens epistrophus idyja aristogiton aristolochiae arthemis arvensis arthemis atratus glaucippe phaethusa philoctetes belti bhagava hippophaes komarovi boisduvali solieri sylvia smaragdinus busiris freija agestis camadeva canens anchisiades gratiosa glaucippe celtis nupta cerialis cerialis aristolochiae chandra nephelus noureddin dispar cippus concolor cerialis hillapana troglodyta cupido pasithoe phlaeas helenus dasarada zerynthia denuba atreus diores domicella dorcas sirtalis coon glaphyra anacardii epicra edusa phlaeas terea oenone oppelii davidii eryx burnii meliboeus euchenor eumeus eunomia palaeno vulgaris solon arcensis falloui feronia impediens februa ferghana februa flegyas thyodamas lativitta histrio buddha glaucippe howqua helenus cippus terea nupta glauconome machaon gratiosa helenor helvina hewitsonii machaon eryx humboldtii ilia helvina hyparete sirtalis sophia iphiclus tyotaroi glaucippe jehana grisea bianor sylvia nevadica philoctetes cippus clyton lucianus euphorbiae klugius luna hyparete macris mars neavei demoleus oregona guatemalena harmodius telamonius orithya chariclea paris sinica glaphyra neavei alala saurita ophione denuba albicinctus haemorrhoidalis azanoides helvina oenone pedemontanus oppelii orbona orbiculare otolais ouida helvina palaeno illioneus parhassus pelaus helenor numilia periboea phaethusa phidippus sylvia phlaeas sirtalis odana anona deidamia populi troglodyta borneensis galathea phlaeas pygela intermedia deva risa fuciformis proximus saurita tithia sambara onolaus saurita ferghana catops saurita nicaea sinaitus splendens sirtalis sophia torosus demoleus acilia burejana styx mathura metis nephelus ferghana orithya arcensis larissa machaon mithridates cyane scabrosus terea polyeuctes terea elegans sirtalis clyton pygas thersites virgata bianor aegeria philoctetes limborgii sondaica philoctetes bomplandii ussheri elegans marcella neonympha bomplandii alida ephesperis xanthus xylina bellona zygia clytia f. clytia clytia f. dissimilis croceus f. helice hypothous subsp. crameri gardetta subsp. darwiniana gardetta subsp. gardetta thrasibulus subsp. thrasibulus

204

, Charles J. Sharp

186

, Benjamin Gimmel

205

, Charles J. Sharp

203

, Andreas Eichler

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